ID   ALLC_RHIME              Reviewed;         340 AA.
AC   Q92VC1;
DT   13-SEP-2005, integrated into UniProtKB/Swiss-Prot.
DT   01-DEC-2001, sequence version 1.
DT   25-MAY-2022, entry version 100.
DE   RecName: Full=Probable allantoicase {ECO:0000255|HAMAP-Rule:MF_00813};
DE            EC=3.5.3.4 {ECO:0000255|HAMAP-Rule:MF_00813};
DE   AltName: Full=Allantoate amidinohydrolase {ECO:0000255|HAMAP-Rule:MF_00813};
GN   Name=alc {ECO:0000255|HAMAP-Rule:MF_00813}; OrderedLocusNames=RB0787;
GN   ORFNames=SMb21283;
OS   Rhizobium meliloti (strain 1021) (Ensifer meliloti) (Sinorhizobium
OS   meliloti).
OG   Plasmid pSymB (megaplasmid 2).
OC   Bacteria; Proteobacteria; Alphaproteobacteria; Hyphomicrobiales;
OC   Rhizobiaceae; Sinorhizobium/Ensifer group; Sinorhizobium.
OX   NCBI_TaxID=266834;
RN   [1]
RP   NUCLEOTIDE SEQUENCE [LARGE SCALE GENOMIC DNA].
RC   STRAIN=1021;
RX   PubMed=11481431; DOI=10.1073/pnas.161294698;
RA   Finan T.M., Weidner S., Wong K., Buhrmester J., Chain P., Vorhoelter F.J.,
RA   Hernandez-Lucas I., Becker A., Cowie A., Gouzy J., Golding B., Puehler A.;
RT   "The complete sequence of the 1,683-kb pSymB megaplasmid from the N2-fixing
RT   endosymbiont Sinorhizobium meliloti.";
RL   Proc. Natl. Acad. Sci. U.S.A. 98:9889-9894(2001).
RN   [2]
RP   NUCLEOTIDE SEQUENCE [LARGE SCALE GENOMIC DNA].
RC   STRAIN=1021;
RX   PubMed=11474104; DOI=10.1126/science.1060966;
RA   Galibert F., Finan T.M., Long S.R., Puehler A., Abola P., Ampe F.,
RA   Barloy-Hubler F., Barnett M.J., Becker A., Boistard P., Bothe G.,
RA   Boutry M., Bowser L., Buhrmester J., Cadieu E., Capela D., Chain P.,
RA   Cowie A., Davis R.W., Dreano S., Federspiel N.A., Fisher R.F., Gloux S.,
RA   Godrie T., Goffeau A., Golding B., Gouzy J., Gurjal M., Hernandez-Lucas I.,
RA   Hong A., Huizar L., Hyman R.W., Jones T., Kahn D., Kahn M.L., Kalman S.,
RA   Keating D.H., Kiss E., Komp C., Lelaure V., Masuy D., Palm C., Peck M.C.,
RA   Pohl T.M., Portetelle D., Purnelle B., Ramsperger U., Surzycki R.,
RA   Thebault P., Vandenbol M., Vorhoelter F.J., Weidner S., Wells D.H.,
RA   Wong K., Yeh K.-C., Batut J.;
RT   "The composite genome of the legume symbiont Sinorhizobium meliloti.";
RL   Science 293:668-672(2001).
CC   -!- CATALYTIC ACTIVITY:
CC       Reaction=allantoate + H2O = (S)-ureidoglycolate + urea;
CC         Xref=Rhea:RHEA:11016, ChEBI:CHEBI:15377, ChEBI:CHEBI:16199,
CC         ChEBI:CHEBI:17536, ChEBI:CHEBI:57296; EC=3.5.3.4;
CC         Evidence={ECO:0000255|HAMAP-Rule:MF_00813};
CC   -!- PATHWAY: Nitrogen metabolism; (S)-allantoin degradation; (S)-
CC       ureidoglycolate from allantoate (aminidohydrolase route): step 1/1.
CC       {ECO:0000255|HAMAP-Rule:MF_00813}.
CC   -!- SIMILARITY: Belongs to the allantoicase family. {ECO:0000255|HAMAP-
CC       Rule:MF_00813}.
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DR   EMBL; AL591985; CAC49187.1; -; Genomic_DNA.
DR   PIR; C95940; C95940.
DR   RefSeq; NP_437327.1; NC_003078.1.
DR   RefSeq; WP_010975643.1; NC_003078.1.
DR   AlphaFoldDB; Q92VC1; -.
DR   SMR; Q92VC1; -.
DR   STRING; 266834.SM_b21283; -.
DR   EnsemblBacteria; CAC49187; CAC49187; SM_b21283.
DR   GeneID; 25012998; -.
DR   GeneID; 61600763; -.
DR   KEGG; sme:SM_b21283; -.
DR   PATRIC; fig|266834.11.peg.5717; -.
DR   eggNOG; COG4266; Bacteria.
DR   HOGENOM; CLU_038797_1_2_5; -.
DR   OMA; MDDGWET; -.
DR   UniPathway; UPA00395; UER00654.
DR   PRO; PR:Q92VC1; -.
DR   Proteomes; UP000001976; Plasmid pSymB.
DR   GO; GO:0004037; F:allantoicase activity; IEA:UniProtKB-UniRule.
DR   GO; GO:0000256; P:allantoin catabolic process; IEA:UniProtKB-UniRule.
DR   GO; GO:0006144; P:purine nucleobase metabolic process; IEA:UniProtKB-KW.
DR   HAMAP; MF_00813; Allantoicase; 1.
DR   InterPro; IPR005164; Allantoicase.
DR   InterPro; IPR015908; Allantoicase_dom.
DR   InterPro; IPR008979; Galactose-bd-like_sf.
DR   PANTHER; PTHR12045; PTHR12045; 1.
DR   Pfam; PF03561; Allantoicase; 2.
DR   PIRSF; PIRSF016516; Allantoicase; 1.
DR   SUPFAM; SSF49785; SSF49785; 2.
DR   TIGRFAMs; TIGR02961; allantoicase; 1.
PE   3: Inferred from homology;
KW   Hydrolase; Plasmid; Purine metabolism; Reference proteome.
FT   CHAIN           1..340
FT                   /note="Probable allantoicase"
FT                   /id="PRO_0000205927"
SQ   SEQUENCE   340 AA;  37135 MW;  FF6A87111810951A CRC64;
     MTRAGEVLPG FAGGTINLAS AGLGARALFA TDEFFGPLER MLKDEPAAFH PGLYDDHGKW
     MDGWETRRRR GAGHDYAVIA LAAKGRIAGF DVDTSHFTGN YPSACSIEAC HSAEDPDEAT
     EWVQLLPVTG LGPNAHHFFA AHSDAVYSHI RLRIHPDGGI ARLRVYGTPA LDLKAMANET
     IDLASCLLGG RIVAFSNGHY GHERLIAPGR GANMGDGWET RRRREPGYDW IIVKLAARGH
     VERILVDTAH FKGNYPDACS LQAADLGGMT AECDMLVASS AMFWNELLPH RKLSADSVHE
     YGSDMLRHAD PVTHVRLNIY PDGGVSRLRI YGRVADPRSR