ID   4CLL9_ORYSJ             Reviewed;         555 AA.
AC   Q7XXL2; Q0JE87;
DT   14-OCT-2008, integrated into UniProtKB/Swiss-Prot.
DT   01-MAR-2004, sequence version 2.
DT   03-AUG-2022, entry version 93.
DE   RecName: Full=4-coumarate--CoA ligase-like 9;
DE            EC=6.2.1.-;
GN   Name=4CLL9; OrderedLocusNames=Os04g0310700, LOC_Os04g24530;
GN   ORFNames=OsJ_013660, OSJNBa0033H08.6;
OS   Oryza sativa subsp. japonica (Rice).
OC   Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Tracheophyta;
OC   Spermatophyta; Magnoliopsida; Liliopsida; Poales; Poaceae; BOP clade;
OC   Oryzoideae; Oryzeae; Oryzinae; Oryza; Oryza sativa.
OX   NCBI_TaxID=39947;
RN   [1]
RP   NUCLEOTIDE SEQUENCE [LARGE SCALE GENOMIC DNA].
RC   STRAIN=cv. Nipponbare;
RX   PubMed=12447439; DOI=10.1038/nature01183;
RA   Feng Q., Zhang Y., Hao P., Wang S., Fu G., Huang Y., Li Y., Zhu J., Liu Y.,
RA   Hu X., Jia P., Zhang Y., Zhao Q., Ying K., Yu S., Tang Y., Weng Q.,
RA   Zhang L., Lu Y., Mu J., Lu Y., Zhang L.S., Yu Z., Fan D., Liu X., Lu T.,
RA   Li C., Wu Y., Sun T., Lei H., Li T., Hu H., Guan J., Wu M., Zhang R.,
RA   Zhou B., Chen Z., Chen L., Jin Z., Wang R., Yin H., Cai Z., Ren S., Lv G.,
RA   Gu W., Zhu G., Tu Y., Jia J., Zhang Y., Chen J., Kang H., Chen X., Shao C.,
RA   Sun Y., Hu Q., Zhang X., Zhang W., Wang L., Ding C., Sheng H., Gu J.,
RA   Chen S., Ni L., Zhu F., Chen W., Lan L., Lai Y., Cheng Z., Gu M., Jiang J.,
RA   Li J., Hong G., Xue Y., Han B.;
RT   "Sequence and analysis of rice chromosome 4.";
RL   Nature 420:316-320(2002).
RN   [2]
RP   NUCLEOTIDE SEQUENCE [LARGE SCALE GENOMIC DNA].
RC   STRAIN=cv. Nipponbare;
RX   PubMed=16100779; DOI=10.1038/nature03895;
RG   International rice genome sequencing project (IRGSP);
RT   "The map-based sequence of the rice genome.";
RL   Nature 436:793-800(2005).
RN   [3]
RP   GENOME REANNOTATION.
RC   STRAIN=cv. Nipponbare;
RX   PubMed=18089549; DOI=10.1093/nar/gkm978;
RG   The rice annotation project (RAP);
RT   "The rice annotation project database (RAP-DB): 2008 update.";
RL   Nucleic Acids Res. 36:D1028-D1033(2008).
RN   [4]
RP   GENOME REANNOTATION.
RC   STRAIN=cv. Nipponbare;
RX   PubMed=24280374; DOI=10.1186/1939-8433-6-4;
RA   Kawahara Y., de la Bastide M., Hamilton J.P., Kanamori H., McCombie W.R.,
RA   Ouyang S., Schwartz D.C., Tanaka T., Wu J., Zhou S., Childs K.L.,
RA   Davidson R.M., Lin H., Quesada-Ocampo L., Vaillancourt B., Sakai H.,
RA   Lee S.S., Kim J., Numa H., Itoh T., Buell C.R., Matsumoto T.;
RT   "Improvement of the Oryza sativa Nipponbare reference genome using next
RT   generation sequence and optical map data.";
RL   Rice 6:4-4(2013).
RN   [5]
RP   NUCLEOTIDE SEQUENCE [LARGE SCALE GENOMIC DNA].
RC   STRAIN=cv. Nipponbare;
RX   PubMed=15685292; DOI=10.1371/journal.pbio.0030038;
RA   Yu J., Wang J., Lin W., Li S., Li H., Zhou J., Ni P., Dong W., Hu S.,
RA   Zeng C., Zhang J., Zhang Y., Li R., Xu Z., Li S., Li X., Zheng H., Cong L.,
RA   Lin L., Yin J., Geng J., Li G., Shi J., Liu J., Lv H., Li J., Wang J.,
RA   Deng Y., Ran L., Shi X., Wang X., Wu Q., Li C., Ren X., Wang J., Wang X.,
RA   Li D., Liu D., Zhang X., Ji Z., Zhao W., Sun Y., Zhang Z., Bao J., Han Y.,
RA   Dong L., Ji J., Chen P., Wu S., Liu J., Xiao Y., Bu D., Tan J., Yang L.,
RA   Ye C., Zhang J., Xu J., Zhou Y., Yu Y., Zhang B., Zhuang S., Wei H.,
RA   Liu B., Lei M., Yu H., Li Y., Xu H., Wei S., He X., Fang L., Zhang Z.,
RA   Zhang Y., Huang X., Su Z., Tong W., Li J., Tong Z., Li S., Ye J., Wang L.,
RA   Fang L., Lei T., Chen C.-S., Chen H.-C., Xu Z., Li H., Huang H., Zhang F.,
RA   Xu H., Li N., Zhao C., Li S., Dong L., Huang Y., Li L., Xi Y., Qi Q.,
RA   Li W., Zhang B., Hu W., Zhang Y., Tian X., Jiao Y., Liang X., Jin J.,
RA   Gao L., Zheng W., Hao B., Liu S.-M., Wang W., Yuan L., Cao M.,
RA   McDermott J., Samudrala R., Wang J., Wong G.K.-S., Yang H.;
RT   "The genomes of Oryza sativa: a history of duplications.";
RL   PLoS Biol. 3:266-281(2005).
RN   [6]
RP   NUCLEOTIDE SEQUENCE [LARGE SCALE MRNA] OF 420-555.
RC   STRAIN=cv. Nipponbare;
RG   The rice full-length cDNA consortium;
RT   "Oryza sativa full length cDNA.";
RL   Submitted (OCT-2006) to the EMBL/GenBank/DDBJ databases.
RN   [7]
RP   GENE FAMILY.
RX   PubMed=18627494; DOI=10.1111/j.1469-8137.2008.02534.x;
RA   de Azevedo Souza C., Barbazuk B., Ralph S.G., Bohlmann J., Hamberger B.,
RA   Douglas C.J.;
RT   "Genome-wide analysis of a land plant-specific acyl:coenzyme A synthetase
RT   (ACS) gene family in Arabidopsis, poplar, rice and Physcomitrella.";
RL   New Phytol. 179:987-1003(2008).
CC   -!- DOMAIN: Both substrate-binding domains (SBD1 and SBD2) are involved in
CC       the substrate recognition, and are sufficient to confer the substrate
CC       specificity.
CC   -!- SIMILARITY: Belongs to the ATP-dependent AMP-binding enzyme family.
CC       {ECO:0000305}.
CC   -!- SEQUENCE CAUTION:
CC       Sequence=BAF14350.1; Type=Erroneous gene model prediction; Evidence={ECO:0000305};
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DR   EMBL; AL662942; CAD37124.3; -; Genomic_DNA.
DR   EMBL; AP008210; BAF14350.1; ALT_SEQ; Genomic_DNA.
DR   EMBL; AP014960; -; NOT_ANNOTATED_CDS; Genomic_DNA.
DR   EMBL; CM000141; EAZ30177.1; -; Genomic_DNA.
DR   EMBL; AK242522; -; NOT_ANNOTATED_CDS; mRNA.
DR   AlphaFoldDB; Q7XXL2; -.
DR   SMR; Q7XXL2; -.
DR   STRING; 4530.OS04T0310800-01; -.
DR   PaxDb; Q7XXL2; -.
DR   PRIDE; Q7XXL2; -.
DR   EnsemblPlants; Os04t0310800-01; Os04t0310800-01; Os04g0310800.
DR   Gramene; Os04t0310800-01; Os04t0310800-01; Os04g0310800.
DR   eggNOG; KOG1176; Eukaryota.
DR   InParanoid; Q7XXL2; -.
DR   PlantReactome; R-OSA-1119316; Phenylpropanoid biosynthesis.
DR   PlantReactome; R-OSA-1119531; Flavonoid biosynthesis.
DR   Proteomes; UP000000763; Chromosome 4.
DR   Proteomes; UP000007752; Chromosome 4.
DR   Proteomes; UP000059680; Chromosome 4.
DR   GO; GO:0005783; C:endoplasmic reticulum; IEA:EnsemblPlants.
DR   GO; GO:0016207; F:4-coumarate-CoA ligase activity; IEA:UniProt.
DR   GO; GO:0005524; F:ATP binding; IEA:UniProtKB-KW.
DR   GO; GO:0016405; F:CoA-ligase activity; IBA:GO_Central.
DR   GO; GO:0004467; F:long-chain fatty acid-CoA ligase activity; IBA:GO_Central.
DR   GO; GO:0031956; F:medium-chain fatty acid-CoA ligase activity; IEA:EnsemblPlants.
DR   GO; GO:0106290; F:trans-cinnamate-CoA ligase activity; IEA:UniProt.
DR   GO; GO:0046949; P:fatty-acyl-CoA biosynthetic process; IBA:GO_Central.
DR   GO; GO:0009698; P:phenylpropanoid metabolic process; IEA:UniProt.
DR   GO; GO:0080110; P:sporopollenin biosynthetic process; IEA:EnsemblPlants.
DR   Gene3D; 3.30.300.30; -; 1.
DR   Gene3D; 3.40.50.12780; -; 1.
DR   InterPro; IPR025110; AMP-bd_C.
DR   InterPro; IPR045851; AMP-bd_C_sf.
DR   InterPro; IPR020845; AMP-binding_CS.
DR   InterPro; IPR000873; AMP-dep_Synth/Lig.
DR   InterPro; IPR042099; ANL_N_sf.
DR   Pfam; PF00501; AMP-binding; 1.
DR   Pfam; PF13193; AMP-binding_C; 1.
DR   PROSITE; PS00455; AMP_BINDING; 1.
PE   2: Evidence at transcript level;
KW   ATP-binding; Ligase; Nucleotide-binding; Reference proteome.
FT   CHAIN           1..555
FT                   /note="4-coumarate--CoA ligase-like 9"
FT                   /id="PRO_0000351635"
FT   REGION          271..342
FT                   /note="SBD1"
FT   REGION          343..410
FT                   /note="SBD2"
FT   CONFLICT        420
FT                   /note="T -> M (in Ref. 6; AK242522)"
FT                   /evidence="ECO:0000305"
SQ   SEQUENCE   555 AA;  59420 MW;  F9CD096E1A4A3E1E CRC64;
     MGDAAVPAMV VEEEEQEHVF RSRFPPVAVP DGVTVPEFVL DGAEAYADRV ALVEAAAGGR
     SYTYGEVARD TARFARALRS VGVRKGHVVV VALPNLAVYP VVSLGIMSAG AVFSGVNPRA
     LAAEIKKQVE DSEAKLVVAN EVAFDKVKDA GVPVIGVGDR ERMPGAISWD GLLAAADRTG
     AGVVPVDAAQ QSDLCALPYS SGTTGVSKGV MLSHRNLVSN LCSSMFAVAP ETAGQVVTLG
     LMPFFHIYGI TGICCATLRH KGTVVVMDRF DLRTFLRALV DHRVMFAPLV PPVMLAMVKS
     PVADEFDLSD LALKSVMTAA APLAPDLLAA FQRKFPGVQV EEAYGLTEHS CITLTHAAGD
     GHGHVAKKSS VGFILPNLEV KFVDPDTGRS LPANTPGELC VRSQSVMQGY YKRKEETERT
     VDGKGWLHTG DVGYIDGDGD VFIVDRIKEL IKYKGFQVAP AELEAVLLSH PSVEDAAVFG
     VPDEEAGEVP VACVVRRHGA EEGEEEIVAY VAERVASYKR VRVLHIVDAI PKSVSGKILR
     RQLRDEFIKR MKPSA