ID   YCF2_ANTAG              Reviewed;        2392 AA.
AC   Q859W7;
DT   10-MAY-2004, integrated into UniProtKB/Swiss-Prot.
DT   01-JUN-2003, sequence version 1.
DT   03-AUG-2022, entry version 64.
DE   RecName: Full=Protein Ycf2 {ECO:0000255|HAMAP-Rule:MF_01330};
GN   Name=ycf2 {ECO:0000255|HAMAP-Rule:MF_01330};
OS   Anthoceros angustus (Hornwort) (Anthoceros formosae).
OG   Plastid; Chloroplast.
OC   Eukaryota; Viridiplantae; Streptophyta; Embryophyta; Anthocerotophyta;
OC   Anthocerotopsida; Anthocerotidae; Anthocerotales; Anthocerotaceae;
OC   Anthoceros.
OX   NCBI_TaxID=48387;
RN   [1]
RP   NUCLEOTIDE SEQUENCE [LARGE SCALE GENOMIC DNA], AND RNA EDITING.
RX   PubMed=12527781; DOI=10.1093/nar/gkg155;
RA   Kugita M., Kaneko A., Yamamoto Y., Takeya Y., Matsumoto T., Yoshinaga K.;
RT   "The complete nucleotide sequence of the hornwort (Anthoceros formosae)
RT   chloroplast genome: insight into the earliest land plants.";
RL   Nucleic Acids Res. 31:716-721(2003).
RN   [2]
RP   NUCLEOTIDE SEQUENCE [MRNA], AND RNA EDITING.
RC   TISSUE=Thallus;
RX   PubMed=12711687; DOI=10.1093/nar/gkg327;
RA   Kugita M., Yamamoto Y., Fujikawa T., Matsumoto T., Yoshinaga K.;
RT   "RNA editing in hornwort chloroplasts makes more than half the genes
RT   functional.";
RL   Nucleic Acids Res. 31:2417-2423(2003).
CC   -!- FUNCTION: Probable ATPase of unknown function. Its presence in a non-
CC       photosynthetic plant (Epifagus virginiana) and experiments in tobacco
CC       indicate that it has an essential function which is probably not
CC       related to photosynthesis.
CC   -!- SUBCELLULAR LOCATION: Plastid, chloroplast stroma {ECO:0000255|HAMAP-
CC       Rule:MF_01330}.
CC   -!- RNA EDITING: Modified_positions=344 {ECO:0000269|PubMed:12527781,
CC       ECO:0000269|PubMed:12711687}, 744 {ECO:0000269|PubMed:12527781,
CC       ECO:0000269|PubMed:12711687}, 861 {ECO:0000269|PubMed:12527781,
CC       ECO:0000269|PubMed:12711687}, 1131 {ECO:0000269|PubMed:12527781,
CC       ECO:0000269|PubMed:12711687}, 1255 {ECO:0000269|PubMed:12527781,
CC       ECO:0000269|PubMed:12711687}, 1687 {ECO:0000269|PubMed:12527781,
CC       ECO:0000269|PubMed:12711687}, 1748 {ECO:0000269|PubMed:12527781,
CC       ECO:0000269|PubMed:12711687}, 1805 {ECO:0000269|PubMed:12527781,
CC       ECO:0000269|PubMed:12711687}, 2158 {ECO:0000269|PubMed:12527781,
CC       ECO:0000269|PubMed:12711687}, 2307 {ECO:0000269|PubMed:12527781,
CC       ECO:0000269|PubMed:12711687}; Note=The nonsense codons at positions
CC       344, 744, 861, 1131, 1255, 1805, 2158 and 2307 are modified to sense
CC       codons.;
CC   -!- SIMILARITY: Belongs to the Ycf2 family. {ECO:0000255|HAMAP-
CC       Rule:MF_01330}.
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DR   EMBL; AB086179; BAC55342.1; -; Genomic_DNA.
DR   EMBL; AB087435; BAC55435.1; -; mRNA.
DR   RefSeq; NP_777406.1; NC_004543.1.
DR   AlphaFoldDB; Q859W7; -.
DR   GeneID; 2553519; -.
DR   GO; GO:0009570; C:chloroplast stroma; IEA:UniProtKB-SubCell.
DR   GO; GO:0005524; F:ATP binding; IEA:UniProtKB-KW.
DR   GO; GO:0016887; F:ATP hydrolysis activity; IEA:InterPro.
DR   Gene3D; 3.40.50.300; -; 1.
DR   HAMAP; MF_01330; Ycf2; 1.
DR   InterPro; IPR003593; AAA+_ATPase.
DR   InterPro; IPR003959; ATPase_AAA_core.
DR   InterPro; IPR027417; P-loop_NTPase.
DR   InterPro; IPR008543; Uncharacterised_Ycf2.
DR   Pfam; PF00004; AAA; 1.
DR   Pfam; PF05695; DUF825; 2.
DR   SMART; SM00382; AAA; 1.
DR   SUPFAM; SSF52540; SSF52540; 1.
PE   2: Evidence at transcript level;
KW   ATP-binding; Chloroplast; Nucleotide-binding; Plastid; RNA editing.
FT   CHAIN           1..2392
FT                   /note="Protein Ycf2"
FT                   /id="PRO_0000223048"
FT   BINDING         1658..1665
FT                   /ligand="ATP"
FT                   /ligand_id="ChEBI:CHEBI:30616"
FT                   /evidence="ECO:0000255|HAMAP-Rule:MF_01330"
SQ   SEQUENCE   2392 AA;  288667 MW;  FA6EF05CCEBD5C9B CRC64;
     MKQKLLENKF SYRKFKLEEI KKYEYLLNSC INWNLIKLVT GIPSNREHLI KLFDLRILSS
     LILRDLRKSE MKKSLILKSF PLLILSMFIH RMNSRNIVEI NNCHLERIIY GGINYREGRD
     EISRRYLHSF MKNFSIPLNY PFSTKKGRER YTNNLLRQKK HIWVFKRNLL GKKYIKPVYD
     KIDFYNFEEW KTLIIKEILP SWKISNQSID KANILLEDKN IEDLKHFFEL YVDDIIRRDY
     HWKNSLDIIS YRDRKNQVNF NLKNNLEFLD KKLFYCLISA FCEKVLSEVE GPFKHKRIKS
     TFNLKNIEDF SDFEVTNKEI FKWELHWWKK KIFQFLDKNH ESDQSIAKSF TFFQNKRFLF
     LENYAEFYTW LLYEDSPFHL KKNKQLLDTA KDTFKEDSFQ LNDRRNQIYS FENKGIFLNI
     LHNFSIYVSN KVRKLNHLKI FSDISIKNSY LINKNLVYHG KGSGGTIYED EKQKTWDTTD
     FYLKREKYFI SKTDFLSRSI EKKCFKSNKY FLDLFLNRGD TNKRSTTESN IHSYDLSSLT
     KAKRKEVKSG SSYEFLEKDS FFFLMKENFV KDFKSITKDF FSNREKKMTD YFPKLINYAF
     LDISSIDESN MSFHITGRHL KDFPLNEWRT NIPSRNYMLS DISRMINLSK IRKKFQNVFF
     VSSISSKRIR QNKNSNDFYL ISLIKLNLSW NRPYLCWSTL YKCNKQYIFN RYLKLKERFM
     RGVEQLNLLI TKPNQNYDKI LYFQVESEIG KFHKSKENEL NYNFFFPLLN GINEDDKFLN
     EMFHKKVEAH SEENFETYIY PTNHRKKLKL WYELNKGSYI CLDNVVKKYY SIYKLKFYLI
     NRMKKLWVER IENGNSSNFI RNIVNGHSSN WKINGREWSD YNIKRDKYIN WNSYIYKWFD
     RTKNLQIFPN WFSDNTSRRW FNEIRFIKPD IFMTYPKKLE KNYYSKVYFI FNTSSNSLLE
     SISKKKNINL RGRVLHNYKK LSRDSEKLIK NKLLPESFLN KNLIKNIIIS LFNNEKNHDI
     FREFPRKTFS LWIYKNRNNG FNSINNFQKN LMMNFHNTNT IEFLDYLHDF HFRYDKRLPF
     FMKEIHIKNY DSTYNKFLRI LPIYSNLRSL SINKIKPFFF QSRIDITLSI QLQVFNDSLS
     NYLGRTCNYI FTSFTNHLYK LLNLLIKINS SIYKETDSCS IQKFLAMTPS GPTILNLKIT
     HYDEFFLGEL TSNFNGYFEF SSEPNLSHTE IQSYKNELLS EFLIEFRNKN NRKLQWIKES
     FLKLNLKENF KYVIGGESID RISINLDFYE KNKNILPYPS LSITDFFDKD DKIKVSKKLY
     FLEKWNFFQN YTSWFFTFEW WRYFSNIPLE TFPEVLLNIT DQSKYISYKT TQYIEEILKD
     LWKNLNFIFQ TNILRKININ SEIRLLKQIN NEQHESIFIY IWSHFQFINL SNAIYLTLIS
     LSVSCFLVSE NYFSTLIGLD YIDSWRRFKV IEYLRDPLRG SYLVERWIYG NQTQIIRTEN
     FFMLLLKNFI HYIKNGRFFL FTRKKLDTWL FHSRTLDLSR RKKDLLVKSV ITERSLSQYR
     LNLNLNHNLR NYDFGYKISE KPGFYYLRYL AETYQKDLVN HSFYSSHLAE KWILLAFWKG
     MISSQKLWQT KILNHESYRI PIPFELDLFS SKGILLVGPT EIGKSYLIKN LAADSYVPLI
     KISISKLLYN KPDVITESWI NILMESLQRL TLILELAKKM SPCIIWIQDI HELNVNRSTE
     NVESDPTFLL GILLKYFHTE FISKSTKGII IIGSTHLPRK VDPALISPNR LDKLMNIRII
     DIFERQEKFS ILLHSKRFYF KNKLLHFNEF AYRTMGYNGR DLVALANEVS LINTTQGNST
     IYIDAVRLAL HRQTLGFTYI NNQMLFCQNE GVLLHKIGKA VIQNIFIKIF PMNPLYIGSD
     LWKKKFYYLS EWFLEPSVTE PTIKELTLLS YILGCLAGLA AGDSWSILEH EPENLIPLNK
     FIENDFDLAC GISESFLVEF SWLEIFQDES INNEIEFFPK IGARHFLNMM QRGISSTTSG
     KFMYKQDGLR NRSIFKKTVQ YKEKLYELAS ETTWAPKIWR LSFIRTKSFD WIGRPNDFES
     LYSFWFLREK EQTFSVSENS REIFHSSQVA QYKTKEELPY ERILSRIRRR NVEELESQLG
     FILSEDQFKI LGFPELSTQY RIEYELFQKP MLFMGGRFIW DPTSLSFETR YSVFSRRELF
     IDEEMRRRLY VTYGARRERE RSRSSQKIKQ FFLRCGYGRD FMNNSSIGWW NGLFLTEKYN
     IEIFKRIEEI SVQLKHPEVF TPVYLYQSWL IENSREKLSR FDLLNHRERW IKLNNKSFHY
     FSIHSTLLES YNYLLNFFIS NRILLDEMTK MLLKDGWLFQ NEIEHFICKM KK